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as you have mcafee installed so you can easily turn windows defender off..go to services.msc buy typing in search in start menu of vista..it will show you services..so open it and find windwos defender service and double click on it and choose disabled..now one more step..use some startup manager..like tune up start up manager and remove windows defender entry from there..now you won't get this error and don't worry you are safe..
if you want to delete it by editing registry without tune up software than go to by typing regedit again in search box of vista it will show many enteries. go to the following:
delete the string value that is:
Windows Defender User Interface
if you from some reason want to revert back than creat the same entry in registry in this location and turn on you service by going to service.msc
you can backup you regisrty value and export the single run key to some where safe place..
Please read this article HERE.
"The 2009 McAfee consumer products for Windows Vista disable Microsoft Windows Defender to improve system performance. Windows Vista users who have installed the 2009 McAfee consumer products should not enable Windows Defender."