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Experieincing this issue as well on ver 5.1.2 on one location. After several packet captures looks like when the machine tries to update the server it is sending a large frame over 1480 Bytes of data and getting dropped. Server communicates file and sends data but machine will not. Looking to see if we can extend the MTU size and try this again. But your problem seems to be different as you mention that it goes away after a reboot.
I believe our MTU size is at the default of 1500 but yes it does stop after a reboot.
We get that error on some of our DE machines as well. When it happens, we just force it to synch again and it works fine. I have always attributed it to WAN traffic getting dirty along the way or the firewall/packet shaper trashing it.